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What is The One Thing?
"Curley" played by Jack Pallance in "City Slickers" holds up his finger. He asks Billy Crystal if he knows what is the secret of life. Curley's finger represents The One Thing: life's most important thing. 
Jason and the Argonauts seek the Golden Fleece. Odysseus  finds The One Thing when, after 20 years, he returns to Ithaca. It is more important than everything that happened in the Odyssey or in the Trojan War. 
The One Thing is the joining of a man and a woman, who, as husband and wife, become The One Thing. They each see The One Thing in the other's eyes when they first discover one another. They feel it in their hearts. The One Thing is celebrated in every love story and in every marriage.
The One Thing is the progenitor, the foundation of human life.